:root{--paper:#fbfaf7;--surface:#fff;--ink:#111619;--ink-soft:#3f494e;--muted:#657076;--relation-muted:#5d686d;--line:#e2e5e3;--line-strong:#ced3d0;--sage:#e9eee6;--sage-ink:#314239;--charcoal:#172126;--gold:#9f8754;--gold-dark:#705b2b;--focus:#296a83;--serif:Georgia, "Times New Roman", Times, serif;--sans:Inter, ui-sans-serif, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;--font-sans:var(--sans);--shell:1440px;--gutter:clamp(1.25rem, 4.5vw, 4.75rem);--section-space:clamp(4.5rem, 8vw, 7.5rem)}*,:before,:after{box-sizing:border-box}html{background:var(--paper);color:var(--ink);font-family:var(--sans);scroll-behavior:smooth;text-rendering:optimizelegibility}body{background:radial-gradient(circle at 85% 5%, #bdb29714, transparent 24rem), var(--paper);min-height:100vh;color:var(--ink);margin:0;font-size:1rem;line-height:1.6;overflow-x:hidden}img,svg{max-width:100%;display:block}a{color:inherit;text-underline-offset:.18em;text-decoration-thickness:.075em}a:focus-visible,summary:focus-visible,button:focus-visible{outline:3px solid var(--focus);outline-offset:4px;border-radius:.2rem}h1,h2,h3,p,dl,dd,ul,ol{margin-top:0}h1,h2,h3{font-family:var(--serif);letter-spacing:-.022em;font-weight:500;line-height:1.12}h1{font-size:clamp(2.75rem,5vw,5.1rem)}h2{font-size:clamp(2rem,3vw,3rem)}h3{font-size:clamp(1.2rem,1.6vw,1.45rem)}em{font-family:var(--serif);font-style:italic}.shell{width:min(100%, var(--shell));padding-inline:var(--gutter);margin-inline:auto}.visually-hidden{clip:rect(0 0 0 0);clip-path:inset(50%);white-space:nowrap;width:1px;height:1px;position:absolute;overflow:hidden}.skip-link{z-index:100;background:var(--charcoal);color:#fff;border-radius:.45rem;padding:.7rem 1rem;font-weight:700;transition:transform .15s;position:fixed;top:.75rem;left:.75rem;transform:translateY(-160%)}.skip-link:focus{transform:translateY(0)}.eyebrow{color:var(--muted);letter-spacing:.14em;text-transform:uppercase;margin-bottom:.7rem;font-size:.75rem;font-weight:750;line-height:1.3}.site-header{z-index:20;border-bottom:1px solid var(--line);backdrop-filter:blur(12px);background:#fbfaf7f0;position:relative}.site-header__inner{justify-content:space-between;align-items:center;gap:2rem;min-height:5.8rem;display:flex}.brand{flex-direction:column;align-items:flex-start;padding-block:.65rem;text-decoration:none;display:inline-flex}.brand__name{font-family:var(--serif);letter-spacing:-.025em;font-size:clamp(1.3rem,2vw,1.7rem);font-weight:600;line-height:1.1}.brand__coverage{color:var(--muted);letter-spacing:.025em;margin-top:.36rem;font-size:.78rem}.desktop-nav ul,.mobile-nav ul{align-items:center;gap:clamp(1.75rem,3.5vw,4rem);margin:0;padding:0;list-style:none;display:flex}.desktop-nav a,.mobile-nav a{min-height:2.75rem;color:var(--ink-soft);align-items:center;font-size:.98rem;font-weight:600;text-decoration:none;display:inline-flex;position:relative}.desktop-nav a:after{content:"";transform-origin:100%;background:currentColor;height:1px;transition:transform .16s;position:absolute;bottom:.2rem;left:0;right:0;transform:scaleX(0)}.desktop-nav a:hover:after,.desktop-nav a[aria-current=page]:after{transform-origin:0;transform:scaleX(1)}.desktop-nav a[aria-current=page]{color:var(--ink)}.mobile-nav{display:none}.review-banner{color:#544b32;background:#f2ecda;border-bottom:1px solid #d8cfae}.review-banner__inner{letter-spacing:.01em;text-align:center;justify-content:center;align-items:center;gap:.55rem;min-height:2.7rem;font-size:.76rem;display:flex}.review-banner__inner strong{color:#312b1d;letter-spacing:.055em;text-transform:uppercase;white-space:nowrap;font-weight:800}.hero{isolation:isolate;border-bottom:1px solid var(--line);background:var(--paper);min-height:clamp(25rem,27vw,28rem);position:relative;overflow:hidden}.hero__media,.hero__media picture,.hero__media img{width:100%;height:100%;position:absolute;inset:0}.hero__media img{object-fit:cover;object-position:right center;width:60%;left:auto;right:0}.hero__media:after{z-index:1;background:linear-gradient(90deg, var(--paper) 0%, var(--paper) 38%, #fbfaf7fc 40%, #fbfaf7f5 41%, #fbfaf7c7 42.5%, #fbfaf785 44%, #fbfaf747 45.5%, #fbfaf71a 47%, transparent 48%);content:"";pointer-events:none;position:absolute;inset:0}.hero__taxon{z-index:2;color:#304239ad;font-family:var(--serif);text-shadow:0 1px #ffffff6b;align-items:baseline;gap:.42rem;margin:0;font-size:.86rem;line-height:1.2;display:flex;position:absolute;top:1.15rem;right:1.3rem}.hero__taxon:before{content:"";background:#485b4f59;width:1.65rem;height:1px}.hero__taxon em{color:#1f3028c7;font-size:1.08em}.hero__taxon span{letter-spacing:.015em;font-size:.88em;font-style:normal}.hero__wash{display:none}.hero__content{z-index:2;min-height:inherit;align-items:center;display:flex;position:relative}.hero__copy{width:min(40%,36rem);padding-block:3.5rem}.hero__credit{z-index:3;color:#314239b8;letter-spacing:.012em;margin:0;font-size:.68rem;font-weight:400;line-height:1.35;position:absolute;bottom:.65rem;right:.85rem}.hero__credit a{color:inherit;text-underline-offset:.16em;text-decoration-color:#31423957}.hero__credit a:hover{color:var(--sage-ink);text-decoration-color:currentColor}.hero h1{max-width:17ch;margin-bottom:1.35rem;font-size:clamp(3rem,4vw,4rem)}.hero__lede{max-width:34rem;color:var(--ink-soft);margin-bottom:2rem;font-size:clamp(1.05rem,1.5vw,1.25rem);line-height:1.55}.hero-search{max-width:34rem;margin-bottom:.8rem}.hero-search label{color:var(--ink-soft);margin-bottom:.45rem;font-size:.78rem;font-weight:750;display:block}.hero-search>div{grid-template-columns:minmax(0,1fr) auto;display:grid}.hero-search input{border:1px solid var(--line-strong);min-width:0;min-height:3rem;color:var(--ink);font:inherit;background:#ffffffe0;border-right:0;border-radius:.55rem 0 0 .55rem;padding:.65rem .9rem}.hero-search .button{border-radius:0 .55rem .55rem 0}.hero__browse{color:var(--ink-soft);font-size:.88rem;font-weight:700}.button{border:1px solid #0000;border-radius:.6rem;justify-content:center;align-items:center;gap:.75rem;min-height:3rem;padding:.65rem 1.2rem;font-size:.95rem;font-weight:700;text-decoration:none;transition:background-color .16s,color .16s,transform .16s;display:inline-flex}.button--primary{background:var(--charcoal);color:#fff}.button--primary:hover{background:#2b393f;transform:translateY(-1px)}.stats{border-bottom:1px solid var(--line);background:#ffffff9e}.stats__inner{grid-template-columns:repeat(4,1fr);display:grid}.stat{min-height:5.25rem;color:inherit;justify-content:flex-start;align-items:center;padding:1rem 1.25rem;text-decoration:none;transition:background-color .16s,box-shadow .16s;display:flex}.stat:hover{background:#ffffffc7}.stat:focus-visible{z-index:1;outline:3px solid var(--gold);outline-offset:-3px;position:relative}.stat:hover .stat__text>span,.stat:focus-visible .stat__text>span{color:var(--ink);text-underline-offset:.2em;text-decoration:underline 1px}.stat+.stat{border-left:1px solid var(--line)}.stat__icon{color:#30393e;flex:0 0 2.75rem;width:2.75rem}.stat__icon svg{fill:none;stroke:currentColor;stroke-linecap:round;stroke-linejoin:round;stroke-width:1.7px}.stat__text{flex-flow:wrap;align-items:baseline;gap:.25rem .55rem;min-width:0;line-height:1.25;display:flex}.stat__text strong{font-family:var(--serif);font-size:1.55rem;font-weight:500;line-height:1}.stat__text>span{color:var(--ink-soft);font-weight:650}.stat__text small{width:100%;color:var(--muted);font-size:.78rem}.section{padding-block:var(--section-space)}.section-heading{margin-bottom:clamp(2rem,4vw,3.5rem)}.section-heading h2,.section-heading p:last-child{margin-bottom:0}.section-heading--with-link{justify-content:space-between;align-items:flex-end;gap:2rem;display:flex}.text-link{align-items:center;gap:.45rem;min-height:2.75rem;font-weight:700;text-decoration:none;display:inline-flex}.text-link:hover{text-decoration:underline}.publication-list{border-top:1px solid var(--line-strong)}.publication-filters{border-top:1px solid var(--line-strong);border-bottom:1px solid var(--line);margin-bottom:1.75rem;padding-block:1.25rem 1rem}.publication-filters__grid{grid-template-columns:minmax(18rem,2fr) repeat(4,minmax(9rem,1fr));align-items:end;gap:1rem;display:grid}.publication-filter{gap:.42rem;display:grid}.publication-filter>span{color:var(--muted);letter-spacing:.09em;text-transform:uppercase;font-size:.72rem;font-weight:750}.publication-filter input,.publication-filter select{border:1px solid var(--line-strong);width:100%;min-height:2.9rem;color:var(--ink);font:inherit;background:#ffffff9e;border-radius:.45rem;padding:.6rem .72rem;font-size:.88rem}.publication-filter input::placeholder{color:#818a8e}.publication-filter input:focus,.publication-filter select:focus{outline:2px solid var(--focus);outline-offset:2px}.publication-filters__summary{justify-content:space-between;align-items:center;gap:1rem;min-height:2rem;margin-top:.85rem;display:flex}.publication-filters__summary>div{min-width:0}.publication-filters__more{border-top:1px solid var(--line);margin-top:1rem;padding-top:.9rem}.publication-filters__more summary{width:fit-content;color:var(--ink-soft);cursor:pointer;font-size:.8rem;font-weight:750}.publication-filters__more-grid{grid-template-columns:repeat(2,minmax(0,1fr));gap:1rem;max-width:42rem;margin-top:1rem;display:grid}.publication-filters__export{color:var(--muted);flex-wrap:wrap;align-items:center;gap:.25rem .7rem;margin-top:.35rem;font-size:.78rem;display:flex}.publication-filters__export-status{min-height:1.2em;color:var(--muted);margin:.25rem 0 0;font-size:.75rem}.publication-filters__summary p{color:var(--muted);margin-bottom:0;font-size:.82rem}.publication-filters__summary button{appearance:none;min-width:2rem;min-height:2rem;color:var(--ink-soft);font:inherit;text-underline-offset:.2em;cursor:pointer;background:0 0;border:0;padding:.3rem .35rem;font-size:.8rem;font-weight:700;text-decoration:underline}.publication-filters__summary button:disabled{color:var(--muted);cursor:default;opacity:.45;text-decoration:none}.publication-filters__summary button:focus-visible{outline:2px solid var(--focus);outline-offset:3px;border-radius:.2rem}.publication-filter-empty{border-top:1px solid var(--line-strong);border-bottom:1px solid var(--line);padding-block:clamp(2.5rem,6vw,5rem)}.publication-filter-empty h3{margin-bottom:.6rem;font-size:clamp(1.6rem,3vw,2.3rem)}.publication-filter-empty>p:last-child{color:var(--muted);margin-bottom:0}.publication-row{border-bottom:1px solid var(--line);grid-template-columns:minmax(7rem,.8fr) minmax(0,5fr) auto;align-items:center;gap:clamp(1rem,3vw,3rem);padding-block:1.75rem;display:grid}.publication-row[hidden]{display:none}.publication-row time{color:var(--muted);align-self:start;font-size:.88rem;font-weight:650}.publication-row h3{max-width:68ch;margin-bottom:.55rem}.publication-row h3,.publication-row__meta,.record>h1,.entity-grid strong{overflow-wrap:anywhere}.publication-row__journal{color:inherit;text-decoration-color:#0000}.publication-row__journal:hover,.publication-row__journal:focus-visible{color:var(--ink);text-decoration-color:currentColor}.publication-row h3 a{text-decoration:none}.publication-row h3 a:hover{text-decoration:underline}.publication-row__meta{color:var(--muted);overflow-wrap:anywhere;flex-wrap:wrap;gap:.2rem .9rem;margin-bottom:0;font-size:.84rem;display:flex}.publication-row__meta span+span:before{color:#a3aaa7;content:"·";margin-right:.9rem}.review-record-flag{color:#5f4a19;background:#f3ead6;border-radius:.35rem;width:fit-content;margin:.55rem 0 0;padding:.28rem .48rem;font-size:.72rem;font-weight:750;line-height:1.25}.publication-row__author{color:inherit;text-underline-offset:.22em;text-decoration-color:#0000;text-decoration-thickness:1px;transition:color .15s,text-decoration-color .15s}.publication-row__author:hover,.publication-row__author:focus-visible{color:var(--ink);text-decoration-color:currentColor}.publication-row__tail{align-items:center;gap:1.4rem;display:flex}.publication-row__external{min-height:2.75rem;color:var(--ink-soft);white-space:nowrap;align-items:center;gap:.35rem;font-size:.78rem;font-weight:750;text-decoration:none;display:inline-flex}.publication-row__external:hover{text-decoration:underline}.badge{background:var(--sage);min-height:2rem;color:var(--sage-ink);border-radius:.45rem;align-items:center;padding:.25rem .7rem;font-size:.78rem;font-weight:700;text-decoration:none;display:inline-flex}a.badge:hover{background:#dfe7dc;text-decoration:underline}.badge--broad{cursor:help;white-space:nowrap}.empty-state{border:1px solid var(--line);background:linear-gradient(110deg, #ffffffeb, #f2f1ebbd), var(--surface);border-radius:.9rem;padding:clamp(2rem,5vw,4.5rem);position:relative;overflow:hidden}.empty-state:after{background:var(--gold);content:"";width:4.5rem;height:1px;position:absolute;top:1.5rem;right:1.5rem}.empty-state h2{max-width:18ch;margin-bottom:.8rem;font-size:clamp(1.7rem,3vw,2.5rem)}.empty-state>p:last-child{max-width:62ch;color:var(--ink-soft);margin-bottom:0}.page-intro{border-bottom:1px solid var(--line);padding-top:clamp(3rem,6vw,5.5rem);padding-bottom:clamp(2.75rem,5vw,4.5rem)}.page-intro h1{max-width:15ch;margin-bottom:1.2rem;font-size:clamp(2.65rem,4.2vw,4.25rem)}.page-intro>p:not(.eyebrow){max-width:66ch;color:var(--ink-soft);margin-bottom:0;font-size:clamp(1.05rem,1.6vw,1.25rem)}.page-intro__facts{grid-template-columns:repeat(3,minmax(0,1fr));gap:1.4rem clamp(1.5rem,4vw,3.25rem);max-width:60rem;margin-top:3rem;margin-bottom:0;display:grid}.author-profiles{border-top:1px solid var(--line-strong);max-width:62rem;margin-top:2.4rem;padding-top:1.2rem}.author-profiles__label{color:var(--muted);letter-spacing:.1em;text-transform:uppercase;margin-bottom:.8rem;font-size:.74rem;font-weight:750}.author-profiles ul{flex-wrap:wrap;gap:.55rem;margin:0;padding:0;list-style:none;display:flex}.author-profiles a{border:1px solid var(--line-strong);min-height:2.65rem;color:var(--ink-soft);background:#ffffff9e;border-radius:999px;align-items:center;gap:.45rem;padding:.55rem .9rem;font-size:.82rem;font-weight:750;text-decoration:none;transition:border-color .15s,background-color .15s,color .15s;display:inline-flex}.author-profiles a:hover{border-color:var(--gold);color:var(--ink);background:#fff}.author-profiles a span{color:var(--muted);font-size:.75rem}.page-intro__facts div{border-top:1px solid var(--line-strong);padding:1rem 0 .25rem}.page-intro__facts div+div{border-left:0;padding-left:0}.page-intro__facts dt{color:var(--muted);letter-spacing:.1em;text-transform:uppercase;font-size:.74rem;font-weight:750}.page-intro__facts dd{font-family:var(--serif);margin:.25rem 0 0;font-size:1.15rem}.taxonomy-root{border-left:2px solid var(--gold);margin-bottom:3rem;padding:.25rem 0 .25rem 1.4rem}.taxonomy-root h2{margin-bottom:.5rem}.taxonomy-root p:last-child{color:var(--muted);margin-bottom:0}.taxon-search{border-bottom:1px solid var(--line);padding-top:clamp(2.5rem,5vw,4rem);padding-bottom:clamp(2.5rem,5vw,4rem)}.taxon-search__heading h2{margin-bottom:1.35rem}.taxon-search__field{gap:.45rem;max-width:36rem;display:grid}.taxon-search__field>span{color:var(--muted);letter-spacing:.1em;text-transform:uppercase;font-size:.72rem;font-weight:750}.taxon-search__field input{border:1px solid var(--line-strong);width:100%;min-height:3rem;color:var(--ink);font:inherit;background:#ffffff9e;border-radius:.45rem;padding:.7rem .85rem}.taxon-search__field input:focus-visible{outline:2px solid var(--focus);outline-offset:2px}.taxon-search__status{min-height:1.3em;color:var(--muted);margin:.65rem 0 0;font-size:.82rem}.taxon-search__results{margin-top:1.5rem}.taxon-tree,.taxon-tree__children{margin:0;padding:0;list-style:none}.taxon-tree{border-top:1px solid var(--line-strong)}.taxon-tree__item{border-bottom:1px solid var(--line)}.taxon-tree__branch summary,.taxon-tree__leaf{grid-template-columns:minmax(7rem,.24fr) minmax(0,1fr) auto;align-items:baseline;gap:1rem;min-width:0;padding:1.15rem .15rem;display:grid}.taxon-tree__branch summary{cursor:pointer;padding-right:2.25rem;list-style:none;position:relative}.taxon-tree__branch summary::-webkit-details-marker{display:none}.taxon-tree__branch summary:before{content:"+";color:var(--muted);font-size:1.25rem;line-height:1;position:absolute;right:.1rem;transform:translateY(.05rem)}.taxon-tree__branch[open]>summary:before{content:"−"}.taxon-tree__branch summary:hover,.taxon-tree__leaf:hover{color:var(--gold-dark)}.taxon-tree__branch summary:focus-visible,.taxon-tree__leaf:focus-visible,.taxon-tree__page-link:focus-visible{outline:2px solid var(--focus);outline-offset:4px}.taxon-tree__leaf{color:inherit;text-decoration:none}.taxon-tree__rank{color:var(--muted);letter-spacing:.12em;text-transform:uppercase;font-size:.7rem;font-weight:750}.taxon-tree strong{min-width:0;font-family:var(--serif);overflow-wrap:anywhere;font-size:clamp(1.12rem,2vw,1.4rem);font-weight:500}.taxon-tree__count{color:var(--muted);white-space:nowrap;padding-right:1.8rem;font-size:.8rem}.taxon-tree__leaf .taxon-tree__count{padding-right:0}.taxon-tree__branch-body{border-left:1px solid var(--line-strong);margin:0 0 1.4rem clamp(1rem,4vw,3.5rem);padding-left:clamp(1rem,3vw,2.5rem)}.taxon-tree__page-link{margin:.2rem 0 .75rem;font-size:.82rem;font-weight:700;display:inline-block}.taxon-tree__group+.taxon-tree__group{margin-top:1.35rem}.taxon-tree__group>h3{color:var(--muted);font-family:var(--sans);letter-spacing:.12em;text-transform:uppercase;margin:.35rem 0 .15rem;font-size:.7rem;font-weight:750}.taxon-tree__children .taxon-tree__item:last-child{border-bottom:0}.entity-grid{grid-template-columns:repeat(3,minmax(0,1fr));gap:1.5rem clamp(1.5rem,3vw,2.6rem);margin:0;padding:0;list-style:none;display:grid}.entity-grid a{border-top:1px solid var(--line-strong);background:0 0;flex-direction:column;justify-content:space-between;min-height:9rem;padding:1.15rem .15rem .8rem;text-decoration:none;transition:border-color .16s,color .16s,transform .16s;display:flex}.entity-grid a:hover{border-color:var(--gold);transform:translateY(-1px)}.entity-grid__rank{color:var(--muted);letter-spacing:.12em;text-transform:uppercase;font-size:.72rem;font-weight:750}.entity-grid strong{font-family:var(--serif);font-size:1.45rem;font-weight:500}.entity-grid a>span:last-child{color:var(--muted);font-size:.82rem}.entity-grid--compact a{min-height:9rem}.author-bibliography-index{padding-bottom:clamp(3rem,6vw,5rem)}.author-bibliography-index__heading,.author-bibliography__heading{max-width:70rem}.author-bibliography-index__heading p,.author-bibliography__heading p{max-width:72ch;color:var(--muted);margin-top:1rem}.author-bibliography-index__grid{grid-template-columns:repeat(2,minmax(0,1fr))}.author-bibliography-index__grid a{min-height:8.5rem}.author-index-all,.author-bibliography{border-top:1px solid var(--line)}.author-index-all__heading{border-bottom:1px solid var(--line-strong);padding-bottom:1.25rem}.author-index-compact{grid-template-columns:repeat(3,minmax(0,1fr));column-gap:clamp(1.5rem,3vw,2.6rem);margin:0;padding:0;list-style:none;display:grid}.author-index-compact__heading{border-top:1px solid var(--line-strong);max-width:70rem;margin-bottom:clamp(1.5rem,3vw,2.25rem);padding-top:clamp(2rem,4vw,3rem)}.author-index-compact__heading h3{margin-bottom:0;font-size:clamp(1.75rem,2.4vw,2.35rem)}.author-index-compact__heading p{max-width:72ch;color:var(--muted);margin:.85rem 0 0}.author-index-compact a{border-top:1px solid var(--line-strong);min-height:4.5rem;font-family:var(--serif);align-items:center;padding:.95rem .15rem;font-size:clamp(1.2rem,1.5vw,1.4rem);font-weight:500;line-height:1.25;text-decoration:none;transition:border-color .16s,color .16s;display:flex}.author-index-compact a:hover{border-color:var(--gold);color:var(--gold-dark)}.author-bibliography__years{border-bottom:1px solid var(--line-strong)}.author-bibliography__year{border-top:1px solid var(--line-strong)}.author-bibliography__year summary{min-height:4.25rem;color:var(--ink);cursor:pointer;justify-content:space-between;align-items:center;gap:1.5rem;padding:.9rem .2rem;list-style:none;display:flex}.author-bibliography__year summary::-webkit-details-marker{display:none}.author-bibliography__year summary:after{width:1.5rem;color:var(--muted);content:"+";text-align:right;flex:0 0 1.5rem;font-size:1.35rem;line-height:1}.author-bibliography__year[open] summary:after{content:"−"}.author-bibliography__year summary>span:first-child{font-family:var(--serif);font-size:clamp(1.55rem,2.5vw,2rem)}.author-bibliography__year summary>span:nth-child(2){color:var(--muted);margin-left:auto;font-size:.8rem;font-weight:700}.author-bibliography__year ol{margin:0;padding:0 0 1.6rem;list-style:none}.author-bibliography__year li{grid-template-columns:minmax(0,1fr) auto;align-items:start;gap:.75rem 2rem;padding:1.15rem .2rem;display:grid}.author-bibliography__year li+li{border-top:1px solid var(--line)}.author-bibliography__citation{min-width:0;color:var(--ink-soft);overflow-wrap:anywhere;margin:0;font-size:.95rem;line-height:1.65}.author-bibliography__contributors{color:var(--ink);font-weight:700}.author-bibliography__title{color:var(--ink)}.author-bibliography__citation a,.author-bibliography__catalogue-link{font-weight:700}.author-bibliography__catalogue-link{min-height:2.75rem;color:var(--ink-soft);white-space:nowrap;align-items:center;gap:.35rem;font-size:.78rem;text-decoration:none;display:inline-flex}.author-bibliography__catalogue-link:hover{text-decoration:underline}.taxon-name--italic{font-family:var(--serif);font-style:italic}.taxon-authority{color:inherit;font-family:var(--serif);font-size:1em;font-style:normal;font-weight:inherit}.breadcrumbs{margin-bottom:2rem}.breadcrumbs ol{color:var(--muted);flex-wrap:wrap;gap:.35rem .75rem;margin:0;padding:0;font-size:.8rem;list-style:none;display:flex}.breadcrumbs li+li:before{content:"/";margin-right:.75rem}.record{padding-top:clamp(2.5rem,5vw,5rem);padding-bottom:var(--section-space)}.record>h1{max-width:26ch;margin-bottom:1.2rem;font-size:clamp(2.15rem,3.8vw,3.75rem)}.record__authors{max-width:70ch;color:var(--ink-soft);margin-bottom:.6rem;font-size:1.1rem}.record__collectives{max-width:82ch;color:var(--muted);margin-bottom:2.25rem;font-size:.86rem;line-height:1.6}.record__authors+.record-summary{margin-top:1.65rem}.record-summary{border-left:2px solid var(--gold);max-width:72rem;margin-bottom:2.5rem;padding:.2rem 0 .2rem 1.4rem}.record-summary h2{font-family:var(--sans);letter-spacing:.1em;text-transform:uppercase;margin-bottom:.65rem;font-size:.76rem;font-weight:800}.record-summary>p{max-width:75ch;color:var(--ink-soft);font-family:var(--serif);margin-bottom:0;font-size:clamp(1.1rem,1.5vw,1.3rem);line-height:1.55}.record-summary>.record-summary__basis{color:var(--muted);font-family:var(--sans);margin-top:.8rem;font-size:.82rem;line-height:1.55}.record-summary__basis strong{color:var(--ink-soft)}.release-note{border-top:1px solid var(--line-strong);max-width:62rem;padding-block:2.5rem}.release-note:first-of-type{border-top:0;padding-top:0}.release-note__meta{color:var(--muted);letter-spacing:.07em;text-transform:uppercase;margin-bottom:.6rem;font-size:.78rem;font-weight:750}.release-note h2{margin-bottom:1.4rem;font-size:clamp(1.8rem,3vw,2.6rem)}.release-note h3{font-family:var(--sans);letter-spacing:.08em;text-transform:uppercase;margin:1.5rem 0 .55rem;font-size:.8rem;font-weight:800}.release-note ul{max-width:72ch;margin-bottom:0;padding-left:1.2rem}.feed-subscription{border-top:1px solid var(--line-strong);max-width:54rem;padding-top:clamp(1.5rem,3vw,2.25rem)}.feed-subscription__intro{max-width:62ch;color:var(--ink-soft);margin-bottom:1.4rem;font-size:clamp(1rem,1.4vw,1.12rem)}.feed-subscription__actions{flex-wrap:wrap;align-items:center;gap:.75rem 1.25rem;display:flex}.feed-subscription__copy{cursor:pointer;min-width:12rem}.feed-subscription__open{color:var(--ink-soft)}.feed-subscription__note,.feed-subscription__status,.feed-subscription__fallback{max-width:62ch;font-size:.82rem}.feed-subscription__note{color:var(--muted);margin:.9rem 0 0}.feed-subscription__status{min-height:1.4em;color:var(--sage-ink);margin:.4rem 0 0;font-weight:700}.feed-subscription__fallback{color:var(--muted);overflow-wrap:anywhere;margin:.75rem 0 0}.record-grid{grid-template-columns:repeat(2,minmax(0,1fr));align-items:start;gap:clamp(2.5rem,5vw,4.5rem);display:grid}.record-panel{border:0;border-top:1px solid var(--line-strong);background:0 0;border-radius:0;padding:clamp(1.5rem,3vw,2.5rem) 0 0}.record-panel h2{margin-bottom:1.75rem;font-size:clamp(1.6rem,2.5vw,2.15rem)}.record-panel--wide{margin-top:clamp(2.5rem,5vw,4.5rem)}.external-records__header{grid-template-columns:minmax(0,1fr) minmax(16rem,.8fr);align-items:end;gap:1.5rem 3rem;margin-bottom:1.75rem;display:grid}.external-records__header h2,.external-records__header p{margin-bottom:0}.external-records__header p{max-width:46ch;color:var(--muted)}.external-records__groups{border-top:1px solid var(--line-strong);grid-template-columns:repeat(2,minmax(0,1fr));display:grid}.external-records__group{border-bottom:1px solid var(--line);min-width:0;padding:1.5rem 1.5rem 1.5rem 0}.external-records__group:nth-child(2n){border-left:1px solid var(--line);padding-left:1.5rem;padding-right:0}.external-records__group-heading{justify-content:space-between;align-items:baseline;gap:1rem;margin-bottom:.85rem;display:flex}.external-records__group-heading h3{font-family:var(--font-sans);letter-spacing:.05em;text-transform:uppercase;margin-bottom:0;font-size:.82rem;font-weight:750}.external-records__group-heading>span{color:var(--muted);font-variant-numeric:tabular-nums;font-size:.78rem}.external-records__list{gap:.75rem;margin:0;padding:0;list-style:none;display:grid}.external-records__list li{border-left:2px solid var(--line-strong);min-width:0;padding-left:.75rem}.external-records__subject{color:var(--ink);margin-bottom:.22rem;display:block}.external-records__identifier{overflow-wrap:anywhere;display:inline}.external-records__identifier code{color:inherit;font-family:ui-monospace,SFMono-Regular,Menlo,Consolas,monospace;font-size:.82rem}.external-records__meta{color:var(--muted);flex-wrap:wrap;gap:.3rem .75rem;margin-top:.28rem;font-size:.75rem;display:flex}.external-records__meta>span+span:before{color:#a3aaa7;content:"·";margin-right:.75rem}.external-records__status{color:var(--sage-ink);font-weight:700}.external-records__status--pending{color:var(--muted);font-weight:600}.related-publications{margin-top:clamp(3.5rem,7vw,6rem)}.related-publications__header{max-width:44rem;margin-bottom:2rem}.related-publications__header h2{margin-bottom:.7rem;font-size:clamp(2rem,4vw,3.25rem)}.related-publications__header>p:last-child{color:var(--muted)}.related-publications__list{border-top:1px solid var(--line-strong);margin:0;padding:0;list-style:none}.related-publications__item{border-bottom:1px solid var(--line);grid-template-columns:minmax(0,1fr);align-items:start;padding-block:1.65rem;display:grid}.related-publications__body h3{max-width:none;margin-bottom:.65rem;font-size:clamp(1.25rem,2vw,1.65rem);line-height:1.15}.related-publications__body h3 a{text-decoration:none}.related-publications__body h3 a:hover,.related-publications__body h3 a:focus-visible{text-underline-offset:.15em;text-decoration:underline}.related-publications__meta{color:var(--muted);flex-wrap:wrap;gap:.2rem .85rem;margin-bottom:0;font-size:.82rem;display:flex}.related-publications__meta>span+span:before{color:#a3aaa7;content:"·";margin-right:.85rem}.metadata-list{margin-bottom:0}.metadata-list div{border-top:1px solid var(--line);grid-template-columns:minmax(8rem,.75fr) minmax(0,1.5fr);gap:1rem;padding-block:.8rem;display:grid}.metadata-list dt{color:var(--muted);letter-spacing:.05em;text-transform:uppercase;font-size:.78rem;font-weight:700}.metadata-list dd{overflow-wrap:anywhere;margin:0}.metadata-note{color:var(--muted);margin-top:.3rem;font-size:.78rem;display:block}.method-note{border-top:1px solid var(--line);color:var(--muted);margin-bottom:clamp(3rem,6vw,5rem);padding-top:1.25rem}.method-note p{max-width:72ch;margin:0}.metadata-links{margin:0;padding:0;list-style:none}.metadata-links li+li{margin-top:.4rem}.metadata-links span{color:var(--muted);font-size:.82rem}.metadata-list dd .taxon-name+.taxon-name:before{content:", "}.citation-tools{border-top:1px solid var(--line);margin-top:1.5rem;padding-top:1.35rem}.citation-tools h3{font-family:var(--font-sans);letter-spacing:.05em;text-transform:uppercase;margin-bottom:.85rem;font-size:.78rem;font-weight:700}.citation-tools__actions{flex-wrap:wrap;gap:.55rem;display:flex}.citation-action{appearance:none;border:1px solid var(--line-strong);color:var(--ink-soft);font:inherit;cursor:pointer;background:0 0;border-radius:.4rem;align-items:center;padding:.55rem .72rem;font-size:.78rem;font-weight:700;line-height:1;text-decoration:none;display:inline-flex}.citation-action:hover{border-color:var(--ink-soft);background:var(--sage);color:var(--ink)}.citation-action:focus-visible{outline:2px solid var(--ink);outline-offset:2px}.citation-tools__status{min-height:1.2em;color:var(--muted);margin:.65rem 0 0;font-size:.75rem}.about-layout{padding-top:var(--section-space);padding-bottom:var(--section-space);gap:clamp(3.5rem,7vw,6rem);display:grid}.about-layout>section{max-width:70rem}.about-layout>section>p:not(.eyebrow),.status-legend>p:not(.eyebrow){max-width:72ch;color:var(--ink-soft)}.status-legend{scroll-margin-top:2rem}.status-legend__list{border-top:1px solid var(--line-strong);grid-template-columns:repeat(2,minmax(0,1fr));margin:2rem 0 0;display:grid}.status-legend__list div{border-bottom:1px solid var(--line);padding:1.15rem 1.25rem 1.15rem 0}.status-legend__list div:nth-child(2n){border-left:1px solid var(--line);padding-left:1.25rem}.status-legend__list dt{font-weight:750}.status-legend__list dd{color:var(--muted);margin-top:.35rem}.update-list{border-top:1px solid var(--line-strong);max-width:68rem;margin:0;padding:0;list-style:none}.update-list li{border-bottom:1px solid var(--line);padding-block:1.5rem}.update-list h3{margin-bottom:.45rem}.update-list p{max-width:68ch;color:var(--muted);margin-bottom:0}.source-list{margin:1.5rem 0 0;padding-left:1.2rem}.source-list li+li{margin-top:.9rem}.source-list p{color:var(--muted);margin-bottom:0}.source-discrepancy{border-left:4px solid var(--gold);background:#f7f2e4;margin-block:1.5rem;padding:1rem 1.2rem}.source-discrepancy p:last-child{margin-bottom:0}.editorial-provenance{grid-template-columns:repeat(2,minmax(0,1fr));gap:1.5rem;margin-top:1.5rem;display:grid}.editorial-provenance>*,.source-discrepancy{overflow-wrap:anywhere;min-width:0}.editorial-provenance h3,.source-list-heading{font-family:var(--sans);letter-spacing:.05em;text-transform:uppercase;margin-bottom:.7rem;font-size:.8rem;font-weight:750}.editorial-provenance ul{margin-bottom:0;padding-left:1.2rem}.source-list-heading{margin-top:1.8rem}.source-list,.source-list li,.source-list p{overflow-wrap:anywhere}.not-found{min-height:65vh;padding-top:clamp(5rem,12vw,10rem);padding-bottom:var(--section-space)}.not-found h1{margin-bottom:1rem}.not-found>p:not(.eyebrow){max-width:48ch;color:var(--ink-soft);margin-bottom:2rem;font-size:1.1rem}.site-footer{border-top:1px solid var(--line);background:#f3f1eb}.site-footer__inner{min-height:8rem;color:var(--muted);justify-content:space-between;align-items:center;gap:2rem;font-size:.78rem;display:flex}.site-footer__inner p{margin-bottom:0}.site-footer__meta{text-align:right;justify-items:end;gap:.6rem;display:grid}.site-footer__meta nav{flex-wrap:wrap;justify-content:flex-end;gap:.35rem 1rem;display:flex}.site-footer__meta a{color:var(--ink-soft);font-weight:700}.site-footer__name{color:var(--ink);font-family:var(--serif);margin-bottom:.2rem;font-size:1.05rem;display:block}@media (width<=980px){.hero__copy{width:min(42%,31rem)}.hero__media img{width:57%}.stats__inner{grid-template-columns:repeat(2,1fr)}.stat:nth-child(3){border-left:0;border-top:1px solid var(--line)}.stat:nth-child(4){border-top:1px solid var(--line)}.entity-grid,.author-index-compact,.publication-filters__grid{grid-template-columns:repeat(2,minmax(0,1fr))}.status-legend__list{grid-template-columns:1fr}.status-legend__list div:nth-child(2n){border-left:0;padding-left:0}.publication-filter--search{grid-column:1/-1}.publication-filters__more-grid{grid-template-columns:1fr}}@media (width<=760px){.editorial-provenance{grid-template-columns:minmax(0,1fr)}:root{--gutter:clamp(1rem, 5vw, 2rem);--section-space:4.5rem}.site-header__inner{gap:1rem;min-height:5rem}.desktop-nav{display:none}.mobile-nav{display:block;position:relative}.mobile-nav summary{border:1px solid var(--line-strong);min-width:3.25rem;min-height:2.75rem;color:var(--ink);cursor:pointer;border-radius:999px;justify-content:center;align-items:center;font-size:.84rem;font-weight:750;list-style:none;display:inline-flex}.mobile-nav summary::-webkit-details-marker{display:none}.mobile-nav[open] summary{background:var(--charcoal);color:#fff}.mobile-nav nav{border:1px solid var(--line);background:var(--surface);border-radius:.75rem;width:min(18rem,100vw - 2rem);padding:.5rem;position:absolute;top:calc(100% + .75rem);right:0;box-shadow:0 1rem 2rem #1116191f}.mobile-nav ul{flex-direction:column;align-items:stretch;gap:0}.mobile-nav a{border-radius:.45rem;width:100%;min-height:3rem;padding-inline:.75rem}.mobile-nav a[aria-current=page]{background:var(--sage)}.review-banner__inner{flex-wrap:wrap;gap:.15rem .45rem;min-height:3.2rem;padding-block:.55rem;line-height:1.35}.hero{background:var(--paper);flex-direction:column;min-height:auto;display:flex}.hero__content{order:1;min-height:auto}.hero__copy{width:100%;padding-block:3.5rem}.hero h1{max-width:12ch;font-size:clamp(2.65rem,12vw,4.1rem)}.hero-search>div{grid-template-columns:1fr;gap:.55rem}.hero-search input,.hero-search .button{border:1px solid var(--line-strong);border-radius:.55rem}.hero__media,.hero__media picture,.hero__media img{position:relative;inset:auto}.hero__media{order:2;height:clamp(15rem,70vw,24rem)}.hero__media picture,.hero__media img{width:100%;height:100%}.hero__media img{object-fit:cover;object-position:center center;width:100%}.hero__media:after{display:none}.hero__taxon{gap:.35rem;font-size:.78rem;top:.9rem;right:.95rem}.hero__taxon:before{width:1.1rem}.section-heading--with-link{flex-direction:column;align-items:flex-start;gap:.75rem}.publication-row{grid-template-columns:1fr;gap:.7rem;padding-block:1.5rem}.publication-row time{font-size:.8rem}.publication-row__tail{justify-content:space-between}.author-bibliography__year li{grid-template-columns:1fr;gap:.35rem}.publication-filters__grid{grid-template-columns:1fr}.publication-filter--search{grid-column:auto}.page-intro__facts,.record-grid,.external-records__header,.external-records__groups{grid-template-columns:1fr}.external-records__group,.external-records__group:nth-child(2n){border-left:0;padding-left:0;padding-right:0}.related-publications__item{grid-template-columns:1fr}.page-intro__facts div+div{border-top:1px solid var(--line-strong);border-left:0;padding-left:0}.metadata-list div{grid-template-columns:1fr;gap:.3rem}.site-footer__inner{flex-direction:column;justify-content:center;align-items:flex-start;gap:1rem;padding-block:2rem}.site-footer__meta{text-align:left;justify-items:start}.site-footer__meta nav{justify-content:flex-start}}@media (width<=520px){.brand__name{font-size:1.16rem}.page-intro h1>.taxon-label{display:block}.page-intro h1>.taxon-label>.taxon-authority{max-width:100%;margin-top:.35rem;font-size:.56em;line-height:1.25;display:block}.brand__coverage{font-size:.7rem}.button{width:100%}.entity-grid,.author-index-compact,.author-bibliography-index__grid{grid-template-columns:1fr}.author-bibliography__year summary{gap:.75rem}.author-bibliography__year summary>span:nth-child(2){text-align:right;max-width:8rem}.taxon-tree__branch summary,.taxon-tree__leaf{grid-template-columns:1fr;gap:.35rem}.taxon-tree__count{white-space:normal;padding-right:0}.taxon-tree__branch-body{margin-left:.45rem;padding-left:.85rem}.stat{justify-content:flex-start;gap:.75rem;min-height:8.25rem;padding-inline:.8rem}.stat__icon{flex-basis:2.2rem;width:2.2rem}.stat__text strong{font-size:1.65rem}.stat__text>span{font-size:.88rem}.stat__text small{font-size:.7rem}.stat:nth-child(3),.stat:nth-child(4){border-top:1px solid var(--line)}.stat:nth-child(odd){border-left:0}.empty-state:after{width:2.5rem}}@media (width<=360px){.page-intro h1>.taxon-label{font-size:.8em}.stats__inner{grid-template-columns:1fr}.stat{min-height:7.5rem;padding-inline:1rem}.stat+.stat{border-top:1px solid var(--line);border-left:0}}@media (prefers-reduced-motion:reduce){html{scroll-behavior:auto}*,:before,:after{scroll-behavior:auto!important;transition-duration:.01ms!important}}@media print{:root{--gutter:0;--section-space:1.5rem}html,body{background:#fff;min-width:0}.site-header,.review-banner,.site-footer,.skip-link,.breadcrumbs,.citation-tools,.related-publications{display:none!important}.record{width:100%;padding:0}.record>h1{max-width:none;font-size:24pt}.record__authors{margin-bottom:.5rem}.record__collectives{margin-bottom:1.5rem}.record-grid{grid-template-columns:1fr}.record-panel{break-inside:avoid;background:#fff;border:0;padding:0}.record-panel+.record-panel,.record-panel--wide{border-top:1px solid #aaa;margin-top:1.5rem;padding-top:1.5rem}}